laphriini blog
 taxonomy and ethology
 of robber flies
 as an
avocation
 
   
  
  About Laphriini Blog
 This blog is intended as an
adjunct to my Laphriini Pages, a website devoted to a tribe of robber flies
(Diptera:  Asilidae), known as the Laphriini.  The tribe includes two described
genera in North America--Laphria Meigen and Lampria Macquart.
Many years ago for my Ph.D. I divided Laphria into four groups of
generic rank, of which I revised two for my dissertation, and partially revised
one of the remaining two.  I never published the revisions for the two groups I
revised, and over the years have sporadically worked toward completing the
revision for the group I almost finished (I know this doesn't sound like I've
done much work post-PhD. but I've probably put in about a year full-time since I
graduated on everything together, including two three-month stretches I spent
working around-the-clock between jobs trying to finish; I think that's a lot,
everything considered).  The purpose of my Laphriini Pages is to make this
material available to the general public, as the covered species include many
large showy flies which should interest collectors and other amateur Dipterists.
Also included is a review of the fourth group, which I did not touch for my
PhD., and initiated after graduating. This incomplete review is the only
material currently posted.  The purpose of the blog is to keep the
Dipterological community informed of the progress of my on-going revision and
review, comment on the process, and generally interact with the taxonomic and
ethological communities at large.
  
  February 3,  2025  --  Retirement
 I retired from my job with the Federal government last week, after 21 years and several months of working there.  Friday, January 31 was my last day.  I'm mentioning this here because for years my plan has been that once I retired I would try to spend more of my time working on robber flies in general and on wrapping up the revisions this blog covers in particular.  I should be posting a lot more soon.  It may take me several weeks to adjust to my new life.  I'm very far behind in everything, and that will all take time to get sorted out.
I turned 73 on January 1, 2025 . . . (read more)
 
  September 10,  2023  --  A Link for Karl (1959)
 I made a mistake in my recent post for August 26.  I said that there was no copy of Karl (1959) on-line.  There is.  The original German version is here.  This web copy is much better than the scan I had in 1978, from which I xeroxed the drawings to
splice in to my translation.  It is also much better quality than the later
copy included in Geller Grimm's CD of asilid literature for research workers.  It includes all the illustrations.  Plates with drawings of several European Laphria
species (Laphria ephippium (Fabr.), L. ignea (Meigen), L. azurea Hermann, L. clavipes Fabr.) are on pages 660 and 661.  For all
four species the basistylus and dististylus are clearly labelled with the abbreviations bst and dst, respecitively.
A copy I converted to DJVU format (@ 600 dpi) is here.  I personally find that . . . (read more)
 
  September 4,  2023  --  Glossary Modified
 After I posted last week I realized that I had never referred to the hypandrium in
either the diagrams in my dissertation or in the illustrations or even in the Glossary included here.
My excuse is that since the hypandrium is small or lacking in the Laphriini it really didn't matter.  Obviously, given what happened,
that was wrong.  I also realized that here at least in the Glossary I had not included definitions of some related terms, namely, the
basistylus, dististylus, shelf of the
basistylus, gonopod, gonocoxite, gonostylus,
and epandrium.  I decided to fix that today.  Each term listed above is now linked to a definition in the Glossary.
I am sure that other terms will eventually have to be added as well.
I also wanted to say more about Ewald Karl's view of the hypandrium in Laphriini, since his is the consensus view at present.  Here is my translation of
the . . . (read more)
 
  August 26,  2023  --  Updated Broken Links
 I finally got around to updating the many broken links on these pages.  Chief among them were the links to photos of type
specimens at the Museum of Comparative Zoology (MCZ).  Not only had the main URL changed, but the unique numbers
had changed as well, and for some even what kind of type they represented.  I also updated the links to other
web pages about robber flies.  Most of the old web page links were out of date.
This is probably a good place to mention the publication of an excellent additional resource for Laphriini from northeastern North America, namely, the Laphria
(Diptera: Asilidae) of Ontario, with a key to the eastern Canadian species of Laphriini and Dasylechia, by Kate G. Lindsay and Stephen A. Marshall, published on the
Canadian Journal of Arthropod Indentification website. As might be expected given the co-author it contains an excellent photographic key to species.  I particularly want to mention
that the depiction of the modifications of the sixth and seventh abdominal segment in the males of one of the groups I identified as a separate genus in my dissertation was
done admirably using photographs--something I had thought would be nearly impossible.
I have only one criticism, and that is, in Figure 3 what is labelled as the "hypandrium" and "shelf of hypandrium" are actually the basistylus and the shelf of the
basistylus, as you can see in the drawing and photograph, here and
here, respectively.
Unfortunately in subsequent references to the male terminalia throughout the paper this mistake is repeated again and again, when the basistyli are referenced.
This would be an easy mistake to make, and . . . (read more)
 
  April 20,  2022  --  Alive and Well
It is hard to believe that it has been seven years since I posted anything here.  That seven years has been a roller-coaster if anything ever was.  At my job, I got promoted; and at home, a varity of changes--including some health issues--made it very difficult to complete any work.  However, I did manage to accomplish something--namely, I was able to purchase and set up everything I will ever possibly need to continue working on robber flies.  Someone once said that when you are old you have plenty of money and no time; whereas, when you are young, the opposite is true.  I can vouch for that.  During that seven years I bought  everything I thought I might ever possibly want, from a phase-contrast microscope to several microphotography setups, as a conscious preparation for when I will have more time to work on asilids when I retire, and, of course, will also have a lot less less discresionary income than at present.  My purpose in this short post is to let everyone know that I have resumed working both on this blog and the associated papers, in other words, what I am supposed to be using all that equipment for, and am alive and fairly well.  I am 70 years old now.  Hopefully I will manage to complete my papers relatively quickly and then work on asilids as a hobby for another ten years.  I plant to retire from work in two years, when I am 72.
I should be attending the North American Dipterists Society (NADS) field meeting in New Jersey June 13-17.
On a technical note, I switched the website server from a local, private company to Amazon Web Services.  For me, this was extremely cheap--AWS hosting is only about one dollar a month.  For you as my audience, it means that you can now bookmark individual pages within the domain, with the bookmark working later to direct you to that exact page, instead of to the index page for the whole site . . . (read more)
 
  July 25,  2015  --  New Laphriini References
I added three refernces (Reichardt
(1929),  Hardy  and   Preece
(1927), and Linsley (1961))  to
the Annotated  Bibliography today.   All three   are quite  old; but   two  were
new to me.    Reichardt  (1929)
was something  I    had,   and  while  working  on   my  PhD.  had    read   and
found indispensible,   but  for  some reason  had forgotten   to  add   both  to
the  Literature  Cited  in    my    dissertation,   and    to    the   Annotated
Bibliography here when   I  began  this     site.   The  other    two  citations
are  apparently  new  to  the  asilid  literature   indices;    they are     not
mentioned          in          Hull
(1962)      or       in      any             of           Lavigne's    three
literature                          updates                                  (Lavigne          (1999),            Lavigne, Dennis,         and
Gowen                  (1978),                   and                      Ghahari,   Lavigne,
and   Geller-Grimm  (2007)).      The   full citations for the three papers
are:
Reichardt,  H.    1929.   Untersuchungen  über  den  Genitalapparat  der
Asiliden.   Zeitschr. wiss.  Zool. 135:257-301 (In
German;    with   detailed    description   of . . .
(read more)
 
    August 10,  2014  --  Miscellaneous Updates and Some New Photos
Since January I've made a lot of small changes in the setup of the Laphriini
Pages that I've been meaning to document.  Most are ongoing parts of more
extensive changes that haven't yet been fully implemented.  For example, I plan
to fix any reference in the bibliography that has an on-line PDF or HTML
version, to show a link to that version.  With this, my bibliography will become
a "one-stop-shop" for North American Laphriini literature.  You will be able to
look at the title, look at the annotation if there is one, or look at the
original article or book, if it is on-line.  I've added links for 59 of the 211
references.  Many of the links are to page readers for large depositories like
Biodiversity Heritage Library and HathiTrust.  Others are to on-line PDFs. When
I've added a link, I've also made it a point to annotate the reference, if it
wasn't annotated already.
The references that now have links to a PDF or HTML version of the original article are:
Adisoemarto (1967)
Alexander (1940)
Andrews (1918)
Back (1904)
Baker and Fischer (1975)
Banks (1911)
Banks (1913)
Banks (1917)
Barnes, Lavers, and Raney (2007)
Bell (1921)
Beutenmüller (1904)
Brimley (1922)
Brimley (1938)
Britton (1920)
Bromley (1914)
Bromley (1924)
Bromley (1931a)
Bromley (1934a)
Bromley (1936)
Bromley (1942)
Bromley (1947)
Bromley (1950a)
Bromley (1950b)
Bromley (1950c)
Bromley (1951)
Brown (1929)
Canden, Suludere, Hasbenli, Cagiran, Lavigne, and Scarbrough (2004)
Cannings (2010)
Cannings (2012)
Cockerell (1927)
Cole and Lovett (1921)
Davis (1910)
Dennis and Lavigne (1976)
Evans (1970)
Greene (1918)
Greene (1910)
Greene (1911)
Harrington (1900)
Hine (1904)
Johnson (1895)
Johnson (1902)
Johnson (1910)
Johnson (1913)
Johnson (1925b)
Johnson (1927)
Knutson (1976)
Lavigne, Barr, and Stephens (2001)
Leonard (1928)
McAtee (1918)
McAtee and Banks (1920)
Nagatomi and Nagatomi (1989)
Quentin (1948)
Rohwer (1912)
Shelly (1979)
Snodgrass (1902)
Thompson (1891)
Walton (1910)
Woodruff (1960)
Young (1910)
As another example, I plan to fix most of the photographs so that you can
click on them to get the original, usually much larger version.  This linked
photo will . . . (read more)
 
    January 15,  2014  --  Another Annotation
I annotated one reference in the Bibliography that was already there, namely
Brimley (1938).  This is one I've been
meaning to get to for a long time.  The complete annotated citation is:
 Brimley, C. S.  1938.  The Insects of North
Carolina, being a list of the insects of North Carolina and their close relatives.
N. C. Dep. Agric., Div. Entomol., 560 pp. (p. 338,
B.[ombomima] affinis Mcq., B. champlainii
Walton, B. cinerea Back, B. divisor Bks.,
B. flavicollis Say, B. grossa Fab., B.
posticata Say, B. sacrator Wlk., B.
thoracica Fab., B. virginica Bks.; L.[ampria]
bicolor Wied., L. rubriventris Mcq.,
L. sp. (near rubriventris but abdomen black); L.[aphria]
aktis McAtee, L. saffrana Fab.,
L. scorpio McAtee, L. sericea Say, L.
sicula McAtee).
As with most of the faunal lists in the Bibliography this one lists localities and a date
range for each species.  At some point I am . . .
(read more)
 
    January 5,  2014  --  A Lone Reference to
Herald in the New Year
I added one annotated reference to the bibliography, namely
Snyder (1957).  The complete
citation is:
 Snyder, K. D.  1957.  Check list of insects of Great
Smoky Mountains National Park.  Privately printed.  Montreat College, Montreat,
NC, 78 pp. (p. 38, Bombomina (Dasyllis)
champlainii Walton, B. divisor Bks., B.
flavicollis Say, B. grossa Fab., B.
sacrator Wlk., Laphria canis Will, L. sericea
Say).
I've also uncovered a few other references but haven't . . .
(read more)
 
    December 7,  2013  --  More References
I added seven annotated references to the bibliography, namely
Felt (1910),
Greene (1910),
Nicolay (1919),
Wandoleck (1905),
Washburn (1905),
Weiss (1915), and
Young (1910).  The complete
references are:
 Felt, E. P.  1910.  Insect types in New York
State Museum, pp. 119-122 IN 25th Report of the State Entomologist
1909.  Education Dept. Bull. 475 (New York State Mus. Bull.
 141), pp. 5-178 (p. 120, Dasyllis cinerea
Back (cotype)).
 Greene, G. M.  1910.  Feldman collecting social.
Entomol. News 21:430-431 (p. 430, Dasyllus
[sic] champlaini Walton).
 Nicolay, A. S. 1919.  Additions to Insects of New
Jersey No. 7.  Entomol. News 30:276-279 (p. 278,
Dasyllis cinerea Back).
 Wandolleck, C.  1905.  Diptera für 1904.
Archiv für Naturgeschichte. 1905(2):787-823 (p. 813,
Dasyllis cinerea n. sp., D. fernaldi
n. sp. ... Back; this is a catalog of all the new Diptera described
in 1904).
 Washburn, F. L.  1905.  Diptera of Minnesota.  Two-winged flies affecting
the farm, garden, stock and household.  Minn. Agr. Expt. Sta. Ann. Rpt.
(1904-1905) 13:19-168, Figs. 4-163, 2 pls. (p. 87, Dasyllis
thoracica, Fab. [also Fig. 79], Laphria sericea, Say [also Plate II,
Fig. 21], and Dasyllis sacrator, Walk. [also Plate II, Fig. 10]).
 Weiss, H. B.  1915.  Additions to Insects of New
Jersey.  Entomol. News 26:101-107 (p. 106,
Dasyllis champlaini Walton).
 Young, D. B.  1910.  Additional list of
Adirondack insects, pp. 123-125 IN 25th Report of the State
Entomologist 1909.  Education Dept. Bull. 475 (New York State
Mus. Bull. 141),  pp. 5-178 (p. 124, Dasyllis
sacrator Walk., Laphria sericea Say).
I also annotated two additional citations . . .
(read more)
 
    November 23,  2013  --  Two More References
I added two annotated references to the bibliography, namely
Beutenmueller (1904) and
Johnson (1913).  The complete references are:
 Beutenmüller, W.  1904.  The types of
Diptera in the collection of the American Museum of Natural History.  Bull.
Amer. Mus. Nat. Hist. 20:87-89 (p. 88, Laphria
astur O. S.).
 Johnson, C.W.  1913.  Insects of Florida.  I.
Diptera.  Bull. Amer. Mus. Nat. Hist. 32(3): 37-90 
(p. 61, Dasyllis grossa Fabr. (D. tergissa Say),
Dasyllis lata, Dasyllis posticata Say, Laphria
saffrana Fabr.).
I also annotated two additional citations that were already present, namely .  . .
(read more)
 
    October 26,  2013  --  Miscellaneous Updates
It's been a while since I last posted.  During the interim I've made a number of small
changes and additions.  I had intended to list them all in one go, here, once they reached a
"critical mass."  Problem is, I didn't write them down, thinking that each was too insignificant
to warrant picking up a pencil.  Now when I want to list them I can only remember a few.
First, I replaced a number of habitus photographs with hopefully better versions.  I replaced
the original blurry analog photo of the female of Laphria flavicollis with a stacked
digital version.  I also replaced the stacked photos I had for both male and female of
Laphria sacrator with new stacked photos having better, more diffuse lighting.
Second, I added several annotated references to the bibliography, namely
Dennis and Barnes (2013) and
Cannings (2010).  The complete references are:
 Dennis, D. S., and J. K. Barnes.  2013.
Pupal cases of four Nearctic species of Laphria (Diptera: Asilidae).
Zootaxa 3681(4):478-492. (L. canis,
L. ferox, L. macquarti, and L.
posticata; key provided to pupal cases of 11 species (including listed
four) of Laphria s. lat. Generic key also provided to
distinguish pupal caes of Laphria s. lat. from those of
Andrenosoma and Lampria [note: Lampria is in the
Laphriini, Andrenosoma. .  . . (read
more)
  
    June  2,  2013  --  Flies Around the House
I ran   a Malaise   trap at   my log   house continuously   from this
last Friday, May 31, through today, Sunday, June 2.  This was the second
time I had  run it  this Spring--the   weekend two  weeks ago  being the
first.  Compared   to last   year I  got  started  later this  one--this
has been  a much colder  and more delayed Spring than normal.
The   reason    I   am    posting  is    that   I    caught   a  male
of Laphria    sacrator.    I've  never   taken   this   species
here before, although I knew it was present in western Pennsylvania.   I
also caught   a  female   Laphria  flavicollis   and  a    male
Laphria aktis.   This   brings   to   eight  the    species  of
Laphriini  I have nabbed just out back  of my house over  the 14 years I
have lived there. They are:
Laphria aktis
Laphria sericea
Laphria index
Laphria scorpio
Laphria winnemana
Laphria sicula
Laphria flavicollis
Laphria sacrator
These eight   species were   all taken   within an  area encomapssing
a circle   about  50   feet in   diamter, in    either a   Malaise trap,
or individually    on    a    pile    of   firewood.     This   firewood
was originally  stacked in  a row  alongside and  somewhat removed  from
my house.  Over the  years  I've let it  rot  to attract insects  rather
than use it in my fireplace.
Six of   these eight   species will  also  eventually  be  placed  in
other genera; however, .  . . (read
more)
  
    May  13,  2013  --  More Literature
I added nine literature citations to the Bibliography.  This nine is a miscellany; only two--Franklin (1912) and Free and Butler (1959)--directly mention North American Laphriini in North America, and the latter is a citation and not an original observation.  The full citations for these two references are:
Franklin, H. J.  1912.    The Bombidae of the
New World.  Trans. Amer. Entomol. Soc. 39:177-486 
(pp. 238, Dasyllis thoracica Fabricius, D. affinis
Macquart, D. tergissa (var.), and D. flavicollis
Say all resemble Bombus impatiens and Psithyrus laboriosus,
and all are found within the ranges of these two bee species; Dasyllis sacrator
Walker resembles Bombus vagans and B. perplexus and is found
within the ranges of these two bee species in the northeastern United States and
southeastern Canada; Dasyllis astur O. S. resembles Bombus californiens
and B. vosnesenskii and it ranges with those species in the Pacific
Coast States; last, Dasyllis posticata Say resembles Bombus fervidus
females and both are distributed widely;  pp. 237-239; discusses Batesian and aggressive
mimicry of bumble bees by robber flies and syrphids.).
Free, J. B. and G. C. Butler.  1959.  Bumble
Bees.  Collins, St. James Place, London.  
(The New Naturalist series; pp. 81-82, Dasyllis grossa).
Three--Wichmann (1956),
Perris (1871), and
Quentin (1948)
--mention the Holarctic species Choerades gilvus in  . . . (read more
)
     May  4,  2013  --  Yet Another Literature
Citation
I         added         another         literature          citation,
namely
Gabritschevsky
(1926).
This reference  isn't  really  new to  me;   it's something I've had   a
copy   of for   a long      time.   I     just      hadn't read it until
this week.  Yet   as far as  I can  tell   it is one  that has     never
been      mentioned       in       any       of      the  various asilid
bibliographies.  It was published  in the Biological Bulletin;    I seem
to   remember  that  that  journal  may   consist  of  translations  of
papers that were first  published in  Russian.   The  paper  itself   is
notable for   species-name     misspellings    as     well    as     for
one species     for    which               the            name        is
completely wrong (Dasyllis dithoracica).     The main  emphasis
is on color convergence  between      bumblebees    and  flies        in
the syrphid   genus    Volucella.         The      way       in
which the thoracic  and       abdominal      patterns  are    diagrammed
for both   flies          and     bees            is         instructive
and possibly eventually    can   serve  as       a        model      for
an updated            analysis                    of               color
patterns within  Laphria    s.          str.        The
full reference  is:
Gabritschevsky,   E.    1926.    Convergence
of  coloration    between
American pilose  flies   and  bumblebees   (Bombus).  Biol.
Bull.. 51(4):269-886.(Divides  American
Laphria     [Dasyllis     at     the      time]    and
Bombus into three   pattern groups,  the first  consisting   of
species in  the eastern United  States, the   second  also  of   eastern
species,    and the     third    of     species   in      the    Rockies
(referred    to  as   the Colorado  area);   p.     273,      the  first
group, consisting  of  . . . (read more
)
  
    March  19,  2013  --  Another Literature
Citation
It's a truism that the more that is happening, the less that is recorded.
It's been that way with me at least for the last several months.  In that time, I've
figured out how to make distribution maps with a free package, Quantum
GIS, that look as nice as the maps I've made at work with ArcGIS
(I've been wanting an affordable package for mapping at home for years).  And
I got myself setup with Corel Draw, Corel Painter, and
AutoDesk SketchBookPro to make line drawings and paintings directly using
a computer, second monitor, and 12x12 Wacom drawing tablet.  And last, I finally
settled on an easy-to-use-and-modify database for documenting and annotating my asilid
collection, called Small Museum Organizer Pro.  I'll post something about each
of these later. For now, though, I'm just going to mention a new literature
citation for Laphria sacrator in Michican, namely Hancock (1911).  The full reference
is:
Hancock, J. L. 1911. Nature Sketches in Temperate
America. xviii + 451 pp., Chapter IV. Mimicry, with Examples (frontispiece of entire book; also p. 127-129, 131 [in subchapter titled
The Bumblebee's Mimic], Dasyllis sacrata [sic].  The model
is stated to be Bombus consimilis, and the observations were made at
Lakeside, Berrien County, Michigan.  Several types of prey are also listed.).
Throughout the book . . . (read more
)
  
    November  18,  2012  --  Removing  an
Unwanted Photo Background
You  may  have  wondered   how   I've  managed  to   clean  up   the
backgrounds so completely    in some   of   my   habitus photos,    such
as    the   three   for   Laphria                       grossa,
here.  The  following is  the method
I use.   Now before  anyone says  anything I  need to  acknowledge  five
things 1) There may be  better ways to do  this in Adobe Photoshop;    I
have an  older version,  5.5.; but   since I  can get  the latest  of GIMP free  hardy use  it  any  more,  2)
Many  of my  older photos  are  extremely crappy --  it  obviously  does
not apply  to   them,   3) Many   of  my newer   photos,    for    which
I      have       used   this   technique,   are    also
extremly   crappy,    4)     So  far    at least    I haven't  used  the
technique everywhere  I could  or probably  should have,  and last, 5) I
obviously still have a lot to learn about photo processing.
The   GIMP  is         free        software;      the         acronym
stands                       for                    "GNU
Image                       Manipulation
Program".        It       is         an      open-source
replacement for  Adobe  Photoshop.    I've    been using  Ver.    2.8.2;
some earlier versions  apparently   do  not   have all   the  resquisite
functions implemented.  You  can, however, do  this with a  much earlier
version,  say for   example  if .  . . (read
more)
  
   November 17, 2012 -- Glossary
Today  I  added a  Glossary of   terms
one  is likely  to find  when perusing  the asilid   literature.  It  is
specifically oriented towards   the older descriptive  literature.  Most
of     the    terms  are  morphological.  The   glossary  itself   is an
amalgam   of   several  public-domain  works,    edited,  modified,  and
tweaked    for  Laphriini.    The tweaking will eventually be extensive.
The glossary  now,  however, includes     much   that     it  shouldn't,
or is  of limited   relevance.   Further,     it    lacks   many  things
that should probably  be  present. These   include many    that  I  know
about,  and many   more    that         I   don't.         Although  the
glossary   is               manifestly             incomplete        and
inadequate,        I       will        be            improving        it
as can.
This  site  much      needs  a       glossary.  For someone      with
a      general    background           neither           the    original
descriptions     nor      my           redescriptions                are
self   explanatory.         You    have            to              start
somewhere. Things    have    a  habit of       never   getting      done
unless  you      put      them    out   for       public    consumption.
But  once   started   they    more-or-less          write    themselves,
bit-by-bit.      This        is      my      philosophy              for
the  entire     website.    With    this   glossary,    eventually    my
mixing,  tweaking,     and      editing of      the   original     works
will    be so         extensive      that   neither    the       overall
result   or  the        individual   entries   will          bear    any
resemblence      to     to                   their              one-time
templates         and props.
Further,  many  of  the  entries  will rely   heavily on my  personal
experience;  dry-as-dust . . . (read more)
 
   November 14, 2012 -- Mistake
Corrected
 Today I removed two references from the Bibliography that I had added recently.  The first
was Aldrich's update to his 1905 Diptera Catalog.  It mentioned a Dasyllis
ablicollis Bigot 1878 from Mexico.  I had not seen this species, and given
the date, thought it might be a Laphria s. str.  I was
wrong.  It is a true Dasyllis, or at least it is treated as one in the
1953 key to that genus by Carrera.  I removed the citation for this paper as
well.  I had yet to see it when I posted, and thought it likely that it would
comment on the confusion between Laphria s. str. and
Dasyllis at the turn of the twentieth century.  I have since looked at
a translation (obtained from Geller-Grimm's website).  It does no
such thing.  So neither paper belongs.  The references for the two papers I
removed are listed . . . (read more)
 
   October 24, 2012 -- More Laphriini
Literature
 Thanks to Bob Lavigne I added 23 new references to North American Laphriini
today.  Bob sent them to me yesterday.  I had known about some of these--more on
that shortly--but some were in places I would have never looked, and had it not
been for Bob would have been missed for a long while if not forever.
 As I said in my last post the current bibliography was derived directly
from the one in my dissertation.  That covered the two genera I plan to describe
as new, as well as the North American Choerades.  In other words it
covered the flies that were once considered as Laphria, before
Bombomima was synonymized, as well as any general papers on
Laphria s. lat., anything of historical significance, and
anything necessary to place a particular species within one of the four genera.
This sounds like a lot.  And in fact it is.  But it is important to note it is
NOT everything, if one wishes exhaustively to cover the entire tribe for
North America north of Mexico.  Namely, it omits references to collection and
prey records for particular species of Laphria s. str.,
if these were segregated from larger works.
 Now I once had all that material and still do.  But it is not in a form I
can easily access.  Back in what seems like dim prehistory when I started my
revision, I indexed all the literature available at the time for everything,
including those disconnected references to particular species of
Laphria s. str.  This was long before the advent of
personal computers.  So my index was in paper form.  It consisted of sheets of 8
1/2 by 11 inch paper, on which I manually typed the citation at the top, and
then either typed (or more usually pasted photocopies) the pertinent lines on
the page below.  I then arranged the sheets in chronological order.  Had they
been amalgamated as typed into a binder it would have been about 10 inches
thick.  The stack was then placed in a small, portable plastic file
cabinet.
 As an aside, I did not originate this method myself.  It was a direct copy
of . . . (read more)
  
   October 20, 2012 -- Laphriini Bibliography
Update
 I updated the Laphriini Bibliography with
13 papers.  To start with, I realized that the bibliography did not include two
essential general asilid papers.  First was the literature update by Lavigne
(1999).  Second was the follow-up update by Ghahari, Lavigne, and Geller-Grimm
(2007).
 Next, I added a newer paper I also knew about but had likewise overlooked,
namely Bedell's (2010) list of Virginia asilids.
 Last, I regularly check the web for new papers.  In doing so I stumbled
across ten much older publications.  Eight mention . . . (read more)
  
   June 14, 2012 -- Early Season Flies
 
 I got out in the woods around my log house on June 9 and took some photos
of live robber flies.  I got shots of two species.  The top one pictured I'm
assuming to be a male of Laphria flavicollis Say.  There were a number
of these about, mostly still hunting in deep forest understory from the upper
surface of fallen timber.  They also occasionally landed on dead leaves on the
forest floor or on live fern fronds.  The one thing in common among all three
types perches was that they were all in bright dappled light, within a foot or
two of the ground.  BTW, the "woods" I'm referring to are old growth oak-maple
forest, with many trees having boles several feet in diameter.
 The next three photos of course aren't laphriines, or even a robber flies.
They're syrphids.  I've included them here because these syrphids were fairly
common around the logs frequented by our L. flavicollis, with
the males in fact apparently patrolling a short beat--in other words, the two
types of fly must regularly come into contact with each other, certainly as
adults and even, now that I know the larvae live in wood, perhaps as immatures.
From looking at Skevington, Thompson, and Marshalls's Field Guide to Syrphidae of the Northeastern North
America, I place them as a species of Temnostoma.  I saw a
L. flavicollis attack one of these patrolling
Temnostoma in mid-air.  I believe it got it, but it flew off before I
could confirm.  These particular Temnostoma wave their front legs
up-and-down when at rest.  This foreleg movement reminds me of the constant
waving by an Ichneumon of it's antennae, but of course otherwise the gestalt of
the two insects differs completely. Supposedly Temnostoma mimic potter
wasps, Eumenidae.  The first two shots were taken several weeks prior, on May 19
(as sort of a displacement activity, when no robber flies were about), and
illustrate well the two most characteristic poses of the fly.  I took the third
shot, of another specimen, a few minutes after the described asilid attack.  It
has more of the specific characters visible, and should be useful if . . . (read more)
  
   May 14, 2012 -- Malaise Trapping,
Miscellaneous Opinions
 This is a laptop with a second screen, set up as it should be to view the
types of plates I will be displaying here.  The plate in the photo is one of
part of the male terminalia of Dasylechia atrox, an oddball Laphriine
fly.  The second monitor is flipped sideways. A setup like this allows you to
take in the entire plate at one glance.  The original lettering on this plate as
well as on most I've done was inked 25-30 years ago with a LeRoy letting set.
Boy, have things changed.  When I made this particular plate computers cost
around ten thousand dollars each and came with all of 64 kilobytes of RAM.  I
remember making this plate well; I did it on Strathsmore board at my home at the
time -- an 8x40 trailer in Laramie, Wyoming, on the edge of town, after work,
with a dissecting scope and grid, light board, and rapidograph set, all of which
I provided myself (and still have).  At the time I worked full-time as a clerk
in a junk store even further away from town but in another direction, and
squeezed in classes and study for my PhD. as I could. In other words, I got
these plates and many others completed under circumstances I'd never want to
repeat.  As for the trailer, I could look out the back window, away from town,
and see nothing but prairie for miles and miles.  I could see so far that
approaching thunderstorms were discrete entities that could be tracked long
before they hit.  In short, you could see those babies coming.
 I sometimes think that graduate students pay too high a price for their
Union Card; considering that once they've gotten a professional job many of them
never do much else again, other than posture and perhaps raise money.  Raising
money, of course is a job too; it takes lots of time and effort, and so I'd
guess the explanation lies there.  But it seems a waste to spend years training
people for some extremely specialized activity like insect taxonomy, for which
presumably they have some natural bent, and then, when they've been hired
professionally, to immediately require as the sine qua non of remaining
employed, that they spend the majority of their time writing grant proposals,
instead of actually exercising and developing their acquired skill, when they
could do the latter almost out of pocket money.  The result is a tiered system
of indentured servants in which nobody wins, lest of all society at large.  I
suppose it's a necessary evil.  At least with funding the average doctoral
candidate gets through in a reasonable period of time, even if he's not doing
what he wants.  And the average professor gets a pay check and security he'd
otherwise be hard put to come by in the rough-and-tumble world of real business.
In fact, generally I'd say grad students are so sick of the whole research gig
by the time they've finished they're glad to be rid of it, like a pennant
exorcising himself of a devil.  I didn't go this route.  But the road I did take
certainly wasn't paved; it had its own ruts and washouts which may in the end
have been far worse. It's a miracle in a way I'm still working on flies, even as
an avocation.  Or I should say, especially as an avocation.
 A week ago I ran a malaise trap in a woodlot I own, next to my log house. A
photo is in the blog header.  It's still very early Spring here (in west central
Pennsylvania), the trees are not far removed from budding, and the ferns are
only a few inches high.  I hadn't run the trap this early before, so I thought
I'd see what I got.  The main catch seemed to be  . . . (read more)
   
  April 22, 2012  -- Cookie Jar
Photography
 I didn't have time this last three weekends to take or process more
photographs of flies.  I did take a couple of shots today of my photography
setup, though.
 The shot just to the right shows the camera mounted on the copy stand from
the side.  I took the lights off the copy stand.  They weren't necessary, and
got in the way when I needed to move it.  The camera is a Nikon D90, the lens is
a Sigma 180 mm macro with a 2x adapter, and the flash is a Sigma ring unit
(EM-140DG). Everything is mounted on a stacking rail.  I've taken all the recent
habitus shots using this setup, with the camera in manual mode, at ASA-200,
F-32, and 1/50th second. I don't think the timing is critical; without the flash
it's dark in the cookie jar; so the shutter just has to be open for longer than
the flash. I set it to 1/30th second for the head profile shots.  One thing that
is very noticeable from this view, is that the camera and lens are not exactly
vertical: they're tilted slightly backwards.  To compensate for this I pin the
fly at a slight angle as well.
 The next photo shows the same rig from an angle, and is slightly enlarged
compared to the first.  You'll note that I'm not using a remote flash; I just
set the timer and then press the button.  My log house is extremely sturdy, but
even so I need to be careful not to move, as with photo stacking any vibration
will result in more work for the software.  BTW, it's the stacking rail that you
adjust incrementally between shots, not the lens focus. Otherwise, the position
of the specimen would shift in the field of view.
 In the third photo, I've shown the pinning tray in the cookie jar.  The
cookie jar does exactly what I need, that is, it diffuses the flash.  But the
pinning tray might be improved on. Other people seem to be pinning the insects
they are photographing on the outside of an inverted Styrofoam cup, and shooting
them from the side, rather than from the top.  Apparently the curved sides of
the cup result in less shadow.  This in turn should result in less work with the
GIMP. I'm considering either using the bottom of the cookie jar itself as a
backdrop or maybe a section of . . . (read
more)
   
  April 7, 2012 -- Musings
  "Down to Gehenna or up to the Throne, He travels fastest who
travels alone."  Rudyard Kipling 
 First off, this isn't a post about Laphriini.  If I had to give it a title,
I would call it "The Importance of Picking the Right Group to Work On."  Or,
more specifically, the right-sized group.  Or something like that, for
as you will see I will wander more than a bit, and would like to keep a tad in
reserve as a surprise.  In short, it's an essay, and if you want to know what I
have to say, you'll need to follow the path where it leads.
 I just turned 60 a few months ago--which is that age where you think a lot
about what you've done and still hope to do.  And in thinking I've realized that
there wouldn't be enough time in five lifetimes to do what I'd originally
thought possible with asilids in just this one, at least without essentially
becoming a machine and boring myself to death.  The Asilidae is a huge group.
There are many North American genera I've never even seen specimens of in a
museum, much less in the field.   Even for the Laphriini, it would have taken me
a lifetime to collect every species I've covered.  And that's just collecting
them as adult flies with a net.  If I'd wanted to collect larvae and rear them
through that alone would take several lifetimes more. 
 I've always had a secret envy of people who in their spare time worked on
small, well-defined groups, with limited distributions.  For example, in the
middle of the last century Mont Cazier worked on apiocerids--a small strange
group of flower-feeding flies that superficially look a little like asilids.  In
his entire life he only published three papers on them--but two were of book
length.  The catch of course is that you have to live where your critters occur;
Cazier spent his summers in Arizona, and apiocerids are distributed throughout
the southwestern deserts.  It's of little use to anybody and especially to
yourself to work on a group that's found so far away you'll be lucky to ever see
a member alive.  You might be able to do this if you work in a museum, but even
so it's a fool's game.  Nothing is really complete or satisfying unless you can
easily relate structure to behavior, behavior to habitat, habitat to
distribution, and all of the previous to some sort of story of . . . (read more)